地点状语能作主语吗
Hominid footprints found in South Africa in the mid-1990s and fossil remains uncovered since 2002 in Central Africa and near the Black Sea suggest that hominids may have walked upright as early as six million years ago.
这里两个and开始并列的都是过去分词作定语修饰的名词短语，但是到第二个and后面突然就变成了near the Black Sea这个地点状语，请问状语作主语该怎么去理解以及有什么规范?
麻烦老师了
最佳答案 2021-10-07 20:53
Hominid footprints found in South Africa in the mid-1990s and fossil remains uncovered since 2002 in Central Africa and near the Black Sea suggest that hominids may have walked upright as early as six million years ago.
两处黑体部分都是过去分词短语作定语。
在 uncovered since 2002 in Central Africa and near the Black Sea 中，黑体部分是uncovered 的地点状语。两个地点状语并列而已：in Central Africa and near the Black Sea
作句子主语的是 Hominid footprints and fossil remains。 near the Black Sea 与in Central Africa 是并列的地点状语，怎么会成了句子的主语？
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