Studies estimated natural community grazing rates by measuring feeding rates of individual zooplankton species in the laboratory and then computing community grazing rates for field conditions using the known population density of grazers.
在句子最后using前面是不是应该加by,因为是通过已知的的密度。
解答如下:仅供参考
Studies【主语】//estimated【谓语动词】// natural community grazing rates【宾语】by measuring feeding rates of individual zooplankton species in the laboratory and then 【介词by省略】computing community grazing rates for field conditions using the known population density of grazers【方式状语】
句子主干: Studies estimated natural community grazing rates.
并列方式状语(1):by measuring feeding rates of individual zooplankton species in the laboratory.
并列方式状语(2):(by) computing community grazing rates for field conditions.
动名词短语:computing community grazing rates for field conditions的方式状语:using the known population density of grazers.
备注(1):using the known population density of grazers为现在分词短语作方式状语修饰动名词短语computing.....for field conditions。
备注(2):并列方式状语(1)和并列方式状语(2)是谓语动词estimated 的方式状语。
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