They suggest a cause of the subsequent state of equilibrium or process
of simplification that would have to have come into operation at some time
after such a pre-historical development.
must have done 和 have to have done 都可以表示根据现在的迹象对过去发生的情况作十分肯定的推测。如果是根据过去的迹象对过去的过去作肯定的推测就不能用情态动词的现在形式,而要用情态动词过去式,所以说话人用了would have to have done 形式表示根据过去的迹象对过去的过去的情况作肯定的推测。