he park is inhabited, and many villages, some of whom the ancestors are dating back to the Angkor period are scattered throughout the park
公园里有人居住,公园里散布着许多村庄,其中一些村庄的祖先可以追溯到吴哥窟时期
请问,whom是连接代词,其先行词是什么?为什么the ancestors are dating back to the Angkor period这句话要用现在进行时?还有,两个逗号的不太明白。