For example, the users of hehe interviewed in The New Yorker article agree on the giggling aspect of hehe, but vary in whether they view it as friendly or conspiratorial: it all depends on how many E’s the word has.

黑体部分为句子的主干：

意思：用户对......看法一致，但在......方面却有分歧。

in 表示“在......方面”，whether 从句作 in 的宾语从句。